Hi Danno,
Welcome to The PainterFactory!
Others may give you better answers to the question of enhancing your brush strokes before printing, but here are a couple of things other Painter artists say they've done that you might try with a test print:
Effects > Focus > Sharpen
or...
Effects > Surface Control > Apply Surface Texture, Using: Image Luminance
The reason you were able to apply Effects > Tonal Control > Equalize to the Layers after they were dropped to the Canvas is that dropping the Layers automatically dries the wet Digital Watercolor paint.
I did a few tests this morning and noticed the result was quite different when I dried the Digital Watercolor without dropping the Layer to the Canvas, then applied Equialize from the result when I first dropped the Layer containing undried Digital Watercolor paint to the Canvas, then applied Equalize.
Test it yourself to see what I mean. You may prefer one method over the other.